IP Library Patent Application 19396666
Patent Application
App. No. 19/396,666

RAS INHIBITORS

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Quick Facts
Patent No.
US None
App. No.
19/396,666
Abstract

The disclosure features macrocyclic compounds, and pharmaceutical compositions and protein complexes thereof, capable of inhibiting Ras proteins, and their uses in the treatment of cancers.

Claims (425)

1 . A compound, or a pharmaceutically acceptable salt thereof, having the structure of Formula I:

wherein:

Q is an optionally substituted 7- to 12-membered bicyclic arylene, an optionally substituted 7- to 12-membered bicyclic heteroarylene, or an optionally substituted 7- to 12-membered bicyclic heterocyclylene, wherein a first ring in Q is bonded to X, and a second ring in Q is bonded to A;

X is a bond; a straight chain C 1 -C 3 alkylene optionally substituted with 1 to 3 substituents independently selected from fluoro, —CN, —C 1 -C 3 alkyl, and —O—C 1 -C 3 alkyl; —O—; —S(O) 0-2 —; *—CH 2 —O—; *—CH 2 —S(O) 0-2 —; *—O—CH 2 —; or *—CH 2 —S(O) 0-2 —, wherein “*” represents a portion of X bound to —C(R 7 )(R 8 )—;

Y is —O—, —NH— or —N(C 1 -C 3 alkyl)-;

R 3 is optionally substituted C 1 -C 6 alkyl, optionally substituted C 3 -C 6 cycloalkyl, optionally substituted C 6 -C 10 aryl, or optionally substituted 3- to 11-membered heterocyclyl;

W is:

wherein:

ring A1 is a 4- to 8-membered cycloalkyl or a 4- to 8-membered heterocyclyl;

W 1 is —N(R 20 )—, —O—, or —C(R 20a )(R 20b )—;

each R A is each independently halo, cyano, hydroxyl, optionally substituted amino, optionally substituted C 1 -C 6 heteroalkyl, or optionally substituted C 1 -C 3 alkyl;

r is 0, 1, 2, 3, or 4;

R 17 , if present, is optionally substituted C 1 -C 6 heteroalkylene or optionally substituted C 1 -C 6 alkylene;

R 18 , if present, is optionally substituted C 1 -C 4 alkylene;

R 19 is optionally substituted C 1 -C 6 heteroalkylene, optionally substituted C 1 -C 6 alkylene, —NH—, or —N(optionally substituted C 1 -C 6 alkyl) or a saturated, nitrogen-containing 3- to 8-membered heterocyclyl;

R 20 is hydrogen or —C 1 -C 3 alkyl;

R 20 is taken together with one R A , the atoms to which they are respectively attached and any intervening atoms, to form an optionally substituted, 5- to 8-membered heterocyclyl that is fused or spiro-fused to ring A, or

R 20 is taken together with any methylene unit in R 18 , or any methylene unit in R 19 , the atoms to which they are respectively attached and any intervening atoms, to form an optionally substituted, 5- to 8-membered heterocyclyl;

each of R 20a and R 20b is, independently, hydrogen, or —C 1 -C 3 alkyl, or R 20a and R 20b are taken together with the carbon atom to which they are bound to form a 3- to 6-membered cycloalkyl ring;

R 16 is O, S, N—CN, or N—O—C 1 -C 3 alkyl;

WH is

each R 22 is, independently, hydrogen, cyano, optionally substituted C 1 -C 3 alkyl, or an optionally substituted 4- to 7-membered saturated heterocyclyl; or R 22 is taken together with either of R 17 or R 19 , the atoms to which they are attached and any intervening atoms to form an optionally substituted 5- to 8-membered ring system;

R 23 is hydrogen, optionally substituted C 1 -C 3 alkyl, or an optionally substituted 4- to 7-membered saturated heterocyclyl;

R 24 is hydrogen, optionally substituted C 1 -C 3 alkyl, or an optionally substituted 4- to 7-membered saturated heterocyclyl; or R 24 is taken together with either of R 17 or R 19 , the atoms to which they are attached and any intervening atoms, to form an optionally substituted 5- to 8-membered ring system;

A is optionally substituted C 2 -C 4 alkylene, optionally substituted C 1 -C 4 heteroalkylene, or optionally substituted C 2 -C 4 alkenylene, optionally substituted 3- to 6-membered cycloalkylene, optionally substituted 3- to 6-membered heterocycloalkylene, optionally substituted 6-membered arylene, or optionally substituted 5- to 10-membered heteroarylene;

L has the structure of Formula VIIa or VIIb:

z is 0, 1, or 2;

X 9 is —NR L6 —, —C(O)—, or —S(O) 2 —;

each of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 is, independently, hydrogen, halogen, hydroxyl, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted C 1 -C 6 heteroalkyl, or optionally substituted C 3 -C 8 cycloalkyl; or any two of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 together with the atoms to which they are attached and any intervening atoms to form an optionally substituted C 3 -C 8 cycloalkyl or a 3- to 8-membered heterocyclyl;

wherein L does not have the structure of

 or

optionally, wherein L is not

R 3 is optionally substituted C 1 -C 6 alkyl, optionally substituted C 3 -C 6 cycloalkyl, optionally substituted C 6 aryl, or optionally substituted 3- to 7-membered heterocyclyl;

R 10 is hydrogen, halogen, optionally substituted C 1 -C 3 alkyl, or C 1 -C 3 optionally substituted heteroalkyl;

R 7 is hydrogen, halogen, or optionally substituted C 1 -C 3 alkyl;

R 8 is hydrogen, halogen, —OH, —CN, optionally substituted C 1 -C 3 heteroalkyl, optionally substituted C 1 -C 3 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted C 6 -C 10 aryl, optionally substituted 4- to 8-membered heteroaryl, optionally substituted C 3 -C 6 cycloalkyl, or optionally substituted 3- to 7-membered heterocyclyl; or

R 7 and R 8 are taken together to form ═CH 2 , an optionally substituted C 3 -C 6 cycloalkyl, or a 3- to 7-membered saturated heterocyclyl; or

R 8 is taken together with a ring atom in Q, the carbon atom to which R 7 is bound and X to form a 4- to 9-membered saturated or unsaturated heterocyclyl that is fused to Q;

R 6 is hydrogen or —CH 3 ;

each R 5 is, independently, halogen, optionally substituted C 1 -C 3 alkyl, or optionally substituted C 1 -C 3 haloalkyl; and

p is 0, 1, 2, or 3, wherein:

(i) the compound is not:

 or

(ii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, two R L1 substituents taken together form spiro cyclopropyl, and W is

 then, Q is not 1-ethyl-indole-2,5-diyl or indole-2,5-diyl substituted with C 1-4 alkyl; or

(iii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, two R L1 substituents taken together form spiro cyclopropyl, and W is

 then, Q is not 1-ethyl-2,5-indol-diyl optionally substituted with 1, 2 or 3 substituents independently selected from halogen, C 1-4 alkyl and C 1-4 alkoxy, wherein the C 1-4 alkyl and C 1-4 alkoxy are each optionally substituted with 1, 2 or 3 substituents independently selected from halogen, OH, C 1-3 alkyl and C 1-3 alkoxy, wherein C 1-3 alkyl and C 1-3 alkoxy are each substituted with 1, 2 or 3 substituents independently selected from deuterium, halogen, OH, methyl, methoxy and OCF 3 ; or

(iv) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, R L1 and R L2 taken together form fused cyclopropyl, and W is

 then Q is not 1-ethyl-2,5-indol-diyl or 2,5-indol-diyl substituted with C 1-4 alkyl; or

(v) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, R L1 and R L2 taken together form fused cyclopropyl, and W is

 then Q is 1-ethyl-2,5-indol-diyl or 2,5-indol-diyl substituted with C 1-4 alkyl; or

(vi) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, R L1 and R L2 taken together form fused cyclopropyl, and W is

 then Q is not 1-ethyl-2,5-indol-diyl optionally substituted with 1, 2 or 3 substituents independently selected from halogen, C 1-4 alkyl and C 1-4 alkoxy, wherein the C 1-4 alkyl and C 1-4 alkoxy are each optionally substituted with 1, 2 or 3 substituents independently selected from halogen, OH, C 1-3 alkyl and C 1-3 alkoxy, wherein C 1-3 alkyl and C 1-3 alkoxy are each substituted with 1, 2 or 3 substituents independently selected from deuterium, halogen, OH, methyl, methoxy and OCF 3 ; or

(vii) when A is -hydroxy-benzene-3,5-diyl or optionally substituted phenylene and two R L1 substituents taken together form spiro cyclopropyl, then W is not

 or

(viii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, R L1 and R L2 taken together form fused cyclopropyl, then W is not

 or

(ix) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, WH is

 and two R L1 substituents taken together form spiro cyclopropyl or spiro cycloalkyl, then A1 is not 1,4-piperazin-diyl optionally substituted with methyl or C 1-4 alkyl; or

(x) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, WH is

 and R L1 and R L2 taken together form fused cyclopropyl or fused cycloalkyl, then A1 is not is not 1,4-piperazin-diyl optionally substituted with methyl or C 1-4 alkyl.

2 . A compound, or a pharmaceutically acceptable salt thereof, having the structure of Formula IIa:

wherein the dotted lines represent zero, one, two, three, or four non-adjacent double bonds;

A is optionally substituted C 2 -C 4 alkylene, optionally substituted C 1 -C 4 heteroalkylene, optionally substituted C 2 -C 4 alkenylene, optionally substituted 3- to 6-membered cycloalkylene, optionally substituted 3- to 6-membered heterocycloalkylene, optionally substituted 6-membered arylene, or optionally substituted 5- to 10-membered heteroarylene;

B is absent, —CH(R 9 )—, >C═CR 9 R 9′ , or >CR 9 R 9′ where the carbon is bound to the carbonyl carbon of —N(R 11 )C(O)—, optionally substituted 3- to 6-membered cycloalkylene, optionally substituted 3- to 6-membered heterocycloalkylene, optionally substituted 6-membered arylene, or optionally substituted 5- to 6-membered heteroarylene;

G is optionally substituted C 1 -C 4 alkylene, optionally substituted C 1 -C 4 alkenylene, optionally substituted C 1 -C 4 heteroalkylene, —C(O)O—CH(R 6 )— where C is bound to —C(R 7 R 8 )—, —C(O)NH—CH(R 6 )—where C is bound to —C(R 7 R 8 )—, optionally substituted C 1 -C 4 heteroalkylene, or 3- to 8-membered heteroarylene;

L 1 is a linker;

W is hydrogen, cyano, S(O) 2 R′, optionally substituted amino, optionally substituted amido, optionally substituted C 1 -C 4 alkoxy, optionally substituted C 1 -C 4 hydroxyalkyl, optionally substituted C 1 -C 4 aminoalkyl, optionally substituted C 1 -C 4 haloalkyl, optionally substituted C 1 -C 4 alkyl, optionally substituted C 1 -C 4 guanidinoalkyl, C 0 -C 4 alkyl optionally substituted 3- to 11-membered heterocycloalkyl, optionally substituted 3- to 8-membered cycloalkyl, or optionally substituted 3- to 8-membered heteroaryl;

L has the structure of Formula VIIa or VIIb:

z is 0, 1, or 2;

X 9 is —NR L6 —, —C(O)—, or —S(O) 2 —;

each of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 is, independently, hydrogen, halogen, hydroxyl, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, or optionally substituted C 1 -C 6 heteroalkyl; or any two of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 together with the atoms to which they are attached and any intervening atoms to form an optionally substituted C 3 -C 8 cycloalkyl or a 3- to 8-membered heterocyclyl;

wherein L does not have the structure of

 or

optionally, wherein L is not

X 1 is optionally substituted C 1 -C 2 alkylene, NR, O, or S(O) q ;

X 2 is O or NH;

X 3 is N or CH;

q is 0, 1, or 2;

R is hydrogen, cyano, optionally substituted C 1 -C 4 alkyl, optionally substituted C 2 -C 4 alkenyl, optionally substituted C 2 -C 4 alkynyl, C(O)R′, C(O)OR′, C(O)N(R′) 2 , S(O)R′, S(O) 2 R′, or S(O) 2 N(R′) 2 ;

each R′ is, independently, hydrogen or optionally substituted C 1 -C 4 alkyl;

Y 1 is C, CH, or N;

Y 2 , Y 3 , Y 4 , and Y 7 are, independently, C or N;

Y 5 is CH, CH 2 , or N;

Y 6 is C(O), CH, CH 2 , or N;

R 13 is cyano, optionally substituted C 1 -C 6 alkyl, optionally substituted C 1 -C 6 heteroalkyl, optionally substituted 3- to 6-membered cycloalkyl, optionally substituted 3- to 6-membered cycloalkenyl, optionally substituted 3- to 6-membered heterocycloalkyl, optionally substituted 6- to 10-membered aryl, or optionally substituted 5- to 10-membered heteroaryl, or

R 13 and R 2 combine with the atoms to which they are attached to form an optionally substituted 3- to 14-membered heterocycloalkyl;

R 2 is absent, hydrogen, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3- to 6-membered cycloalkyl, optionally substituted 3- to 14-membered heterocycloalkyl, optionally substituted 6-membered aryl, optionally substituted 5- to 10-membered heteroaryl;

R 14 is absent or R 2 and R 14 combine with the atom to which they are attached to form an optionally substituted 3- to 8-membered cycloalkyl or optionally substituted 3- to 14-membered heterocycloalkyl;

R 15 is absent, hydrogen, halogen, cyano, or methyl optionally substituted with 1 to 3 halogens;

R 5 is hydrogen, C 1 -C 4 alkyl optionally substituted with halogen, cyano, hydroxy, or C 1 -C 4 heteroalkyl, cyclopropyl, or cyclobutyl;

R 6 is hydrogen or methyl; R 7 is hydrogen, halogen, or optionally substituted C 1 -C 3 alkyl, or

R 6 and R 7 combine with the carbon atoms to which they are attached to form an optionally substituted 3- to 6-membered cycloalkyl or optionally substituted 3- to 7-membered heterocycloalkyl;

R 8 is hydrogen, halogen, hydroxy, cyano, optionally substituted C 1 -C 3 heteroalkyl, optionally substituted C 1 -C 3 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3- to 8-membered cycloalkyl, optionally substituted 3- to 14-membered heterocycloalkyl, optionally substituted 5- to 10-membered heteroaryl, or optionally substituted 6- to 10-membered aryl, or

R 7 and R 8 combine with the carbon atom to which they are attached to form C═CR 7′ R 8′ ; C═N(OH), C═N(O—C 1 -C 3 alkyl), C═O, C═S, C═NH, optionally substituted 3- to 6-membered cycloalkyl, or optionally substituted 3- to 7-membered heterocycloalkyl;

R 7a and R 8a are, independently, hydrogen, halogen, optionally substituted C 1 -C 3 alkyl, or combine with the carbon to which they are attached to form a carbonyl;

R 7′ is hydrogen, halogen, or optionally substituted C 1 -C 3 alkyl;

R 8′ is hydrogen, halogen, hydroxy, cyano, optionally substituted C 1 -C 3 heteroalkyl, optionally substituted C 1 -C 3 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3- to 8-membered cycloalkyl, optionally substituted 3- to 14-membered heterocycloalkyl, optionally substituted 5- to 10-membered heteroaryl, or optionally substituted 6- to 10-membered aryl, or

R 7′ and R 8′ combine with the carbon atom to which they are attached to form optionally substituted 3- to 6-membered cycloalkyl or optionally substituted 3- to 7-membered heterocycloalkyl;

R 9 is hydrogen, fluoro, optionally substituted C 1 -C 6 alkyl, optionally substituted C 1 -C 6 heteroalkyl, optionally substituted 3- to 6-membered cycloalkyl, or optionally substituted 3- to 7-membered heterocycloalkyl; or

R 9 and L combine with the atoms to which they are attached to form an optionally substituted 3- to 14-membered heterocycloalkyl;

R 9′ is hydrogen or optionally substituted C 1 -C 6 alkyl; or

R 9 and R 9′ , combined with the atoms to which they are attached, form a 3- to 6-membered cycloalkyl or a 3- to 6-membered heterocycloalkyl;

R 10 is hydrogen, halogen, hydroxy, optionally substituted C 1 -C 3 heteroalkyl, or optionally substituted C 1 -C 3 alkyl;

R 10a is hydrogen or halogen;

R 11 is hydrogen or optionally substituted C 1 -C 3 alkyl; and

R 21 is hydrogen or optionally substituted C 1 -C 3 alkyl wherein:

(i) the compound is not

 or

(ii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, two R L1 substituents taken together form spiro cyclopropyl, L 1 is —N(CH 3 )C(O)— or —N(CH 3 )C(O)CH 2 N(CH 3 )— and B is —CH(R 9 )—, wherein R 9 is isopropyl or cyclopentyl, then W is not 3-cyclopropylaziridin-2-yl, 3-phenyltetrahydrofuran-2-yl, optionally substituted aziridin-2-yl or optionally substituted tetrahydrofuran-2-yl; or

(iii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, two R L1 substituents taken together form spiro cyclopropyl, L 1 is —N(CH 3 )C(O)— or —N(CH 3 )C(O)CH 2 N(CH 3 )— and B is —CH(R 9 )—, wherein R 9 is C 1-6 alkyl or 3- to 6-membered cycloalkyl, then W is not 3-cyclopropylaziridin-2-yl, 3-phenyltetrahydrofuran-2-yl, optionally substituted aziridin-2-yl or optionally substituted tetrahydrofuran-2-yl; or

(iv) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, R L1 and R L2 taken together form optionally substituted fused cyclopropyl, L 1 is —N(CH 3 )C(O)— or —N(CH 3 )C(O)CH 2 N(CH 3 )— and B is —CH(R 9 )—, wherein R 9 is isopropyl or cyclopentyl, then W is not 3-cyclopropylaziridin-2-yl, 3-phenyltetrahydrofuran-2-yl, optionally substituted aziridin-2-yl or optionally substituted tetrahydrofuran-2-yl; or

(v) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, R L1 and R L2 taken together form optionally substituted fused cyclopropyl, L 1 is —N(CH 3 )C(O)— or —N(CH 3 )C(O)CH 2 N(CH 3 )— and B is —CH(R 9 )—, wherein R 9 is C 1-6 alkyl or 3- to 6-membered cycloalkyl, then W is not 3-cyclopropylaziridin-2-yl, 3-phenyltetrahydrofuran-2-yl, optionally substituted aziridin-2-yl or optionally substituted tetrahydrofuran-2-yl; or

(vi) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, two R L1 substituents taken together form optionally substituted spiro cyclopropyl, L 1 is —N(CH 3 )C(O)— or —N(CH 3 )C(O)CH 2 N(CH 3 )—, R 14 is absent and B is —CH(R 9 )—, wherein R 9 is isopropyl or cyclopentyl, then R 2 is not ethyl or optionally substituted C 1-6 alkyl; or

(vii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, R L1 and R L2 taken together form optionally substituted fused cyclopropyl, L 1 is —N(CH 3 )C(O)— or —N(CH 3 )C(O)CH 2 N(CH 3 )—, R 14 is absent and B is —CH(R 9 )—, wherein R 9 is isopropyl or cyclopentyl, then R 2 is not ethyl or optionally substituted C 1-6 alkyl; or

(viii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene and two R L1 substituents taken together form optionally substituted spiro cyclopropyl, then R 2 is optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3- to 6-membered cycloalkyl, optionally substituted 3- to 14-membered heterocycloalkyl, optionally substituted 6-membered aryl, optionally substituted 5- to 10-membered heteroaryl; or

(ix) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, R L1 and R L2 taken together form optionally substituted fused cyclopropyl, then R 2 is optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3- to 6-membered cycloalkyl, optionally substituted 3- to 14-membered heterocycloalkyl, optionally substituted 6-membered aryl, optionally substituted 5- to 10-membered heteroaryl; or

(x) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene and two R L1 substituents taken together form optionally substituted spiro cyclopropyl, then R 2 and R 14 combine with the atom to which they are attached to form an optionally substituted 3- to 8-membered cycloalkyl or optionally substituted 3- to 14-membered heterocycloalkyl; or

(xi) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, R L1 and R L2 taken together form optionally substituted fused cyclopropyl, then R 2 is optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3- to 6-membered cycloalkyl, optionally substituted 3- to 14-membered heterocycloalkyl, optionally substituted 6-membered aryl, optionally substituted 5- to 10-membered heteroaryl.

3 . A compound, or a pharmaceutically acceptable salt thereof, having the structure of Formula IIb:

wherein the dotted lines represent zero, one, two, three, or four non-adjacent double bonds;

A is optionally substituted C 2 -C 4 alkylene, optionally substituted C 1 -C 4 heteroalkylene, optionally substituted C 2 -C 4 alkenylene, optionally substituted 3- to 6-membered cycloalkylene, optionally substituted 3- to 6-membered heterocycloalkylene, optionally substituted 6-membered arylene, or optionally substituted 5- to 10-membered heteroarylene;

B is absent, —CH(R 9 )—, >C═CR 9 R 9′ , or >CR 9 R 9′ where the carbon is bound to the carbonyl carbon of —N(R 11 )C(O)—, optionally substituted 3- to 6-membered cycloalkylene, optionally substituted 3- to 6-membered heterocycloalkylene, optionally substituted 6-membered arylene, or optionally substituted 5- to 6-membered heteroarylene;

G is optionally substituted C 1 -C 4 alkylene, optionally substituted C 1 -C 4 alkenylene, optionally substituted C 1 -C 4 heteroalkylene, —C(O)O—CH(R 6 )— where C is bound to —C(R 7 R 8 )—, —C(O)NH—CH(R 6 )—where C is bound to —C(R 7 R 8 )—, optionally substituted C 1 -C 4 heteroalkylene, or 3- to 8-membered heteroarylene;

L 1 is a linker;

W is a cross-linking group comprising a carbodiimide, an oxazoline, a thiazoline, a chloroethyl urea, a chloroethyl thiourea, a chloroethyl carbamate, a chloroethyl thiocarbamate, an aziridine, a trifluoromethyl ketone, a boronic acid, a boronic ester, an N-ethoxycarbonyl-2-ethoxy-1,2-dihydroquinoline (EEDQ), an iso-EEDQ or other EEDQ derivative, an epoxide, an oxazolium, or a glycal;

L has the structure of Formula VIIa or VIIb:

z is 0, 1, or 2;

X 9 is —NR L6 —, —C(O)—, or —S(O) 2 —;

each of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 is, independently, hydrogen, halogen, hydroxyl, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, or optionally substituted C 1 -C 6 heteroalkyl; or any two of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 together with the atoms to which they are attached and any intervening atoms to form an optionally substituted C 3 -C 8 cycloalkyl or a 3- to 8-membered heterocyclyl;

wherein L does not have the structure of

 or

optionally, wherein L is not

X 1 is optionally substituted C 1 -C 2 alkylene, NR, O, or S(O) q ;

X 2 is O or NH;

X 3 is N or CH;

q is 0, 1, or 2;

R is hydrogen, cyano, optionally substituted C 1 -C 4 alkyl, optionally substituted C 2 -C 4 alkenyl, optionally substituted C 2 -C 4 alkynyl, C(O)R′, C(O)OR′, C(O)N(R′) 2 , S(O)R′, S(O) 2 R′, or S(O) 2 N(R′) 2 ;

each R′ is, independently, hydrogen or optionally substituted C 1 -C 4 alkyl;

Y 1 is C, CH, or N;

Y 2 , Y 3 , Y 4 , and Y 7 are, independently, C or N;

Y 5 is CH, CH 2 , or N;

Y 6 is C(O), CH, CH 2 , or N;

R 13 is cyano, optionally substituted C 1 -C 6 alkyl, optionally substituted C 1 -C 6 heteroalkyl, optionally substituted 3- to 6-membered cycloalkyl, optionally substituted 3- to 6-membered cycloalkenyl, optionally substituted 3- to 6-membered heterocycloalkyl, optionally substituted 6- to 10-membered aryl, or optionally substituted 5- to 10-membered heteroaryl, or

R 13 and R 2 combine with the atoms to which they are attached to form an optionally substituted 3- to 14-membered heterocycloalkyl;

R 2 is absent, hydrogen, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3- to 6-membered cycloalkyl, optionally substituted 3- to 7-membered heterocycloalkyl, optionally substituted 6-membered aryl, optionally substituted 5- or 6-membered heteroaryl; R 14 is absent or R 2 and R 14 combine with the atom to which they are attached to form an optionally substituted 3- to 8-membered cycloalkyl or optionally substituted 3- to 14-membered heterocycloalkyl;

R 15 is absent, hydrogen, halogen, cyano, or methyl optionally substituted with 1 to 3 halogens;

R 5 is hydrogen, C 1 -C 4 alkyl optionally substituted with halogen, cyano, hydroxy, or C 1 -C 4 heteroalkyl, cyclopropyl, or cyclobutyl;

R 6 is hydrogen or methyl; R 7 is hydrogen, halogen, or optionally substituted C 1 -C 3 alkyl, or

R 6 and R 7 combine with the carbon atoms to which they are attached to form an optionally substituted 3- to 6-membered cycloalkyl or optionally substituted 3- to 7-membered heterocycloalkyl;

R 8 is hydrogen, halogen, hydroxy, cyano, optionally substituted C 1 -C 3 heteroalkyl, optionally substituted C 1 -C 3 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3- to 8-membered cycloalkyl, optionally substituted 3- to 14-membered heterocycloalkyl, optionally substituted 5- to 10-membered heteroaryl, or optionally substituted 6- to 10-membered aryl, or

R 7 and R 8 combine with the carbon atom to which they are attached to form C═CR 7′ R 8′ ; C═N(OH), C═N(O—C 1 -C 3 alkyl), C═O, C═S, C═NH, optionally substituted 3- to 6-membered cycloalkyl, or optionally substituted 3- to 7-membered heterocycloalkyl;

R 7a and R 8a are, independently, hydrogen, halogen, optionally substituted C 1 -C 3 alkyl, or combine with the carbon to which they are attached to form a carbonyl;

R 7′ is hydrogen, halogen, or optionally substituted C 1 -C 3 alkyl; R 8′ is hydrogen, halogen, hydroxy, cyano, optionally substituted C 1 -C 3 heteroalkyl, optionally substituted C 1 -C 3 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3- to 8-membered cycloalkyl, optionally substituted 3- to 14-membered heterocycloalkyl, optionally substituted 5- to 10-membered heteroaryl, or optionally substituted 6- to 10-membered aryl, or

R 7′ and R 8′ combine with the carbon atom to which they are attached to form optionally substituted 3- to 6-membered cycloalkyl or optionally substituted 3- to 7-membered heterocycloalkyl;

R 9 is hydrogen, fluoro, optionally substituted C 1 -C 6 alkyl, optionally substituted C 1 -C 6 heteroalkyl, optionally substituted 3- to 6-membered cycloalkyl, or optionally substituted 3- to 7-membered heterocycloalkyl; or

R 9 and L combine with the atoms to which they are attached to form an optionally substituted 3- to 14-membered heterocycloalkyl;

R 9′ is hydrogen or optionally substituted C 1 -C 6 alkyl; or

R 9 and R 9′ , combined with the atoms to which they are attached, form a 3- to 6-membered cycloalkyl or a 3- to 6-membered heterocycloalkyl;

R 10 is hydrogen, halogen, hydroxy, optionally substituted C 1 -C 3 heteroalkyl, or optionally substituted C 1 -C 3 alkyl;

R 10a is hydrogen or halogen;

R 11 is hydrogen or optionally substituted C 1 -C 3 alkyl; and

R 21 is hydrogen or optionally substituted C 1 -C 3 alkyl wherein:

(i) the compound is not

 or

(ii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, W is 3-cyclopropylaziridin-2-yl or optionally substituted aziridine and two R L1 substituents taken together form spiro cyclopropyl, then R 2 is not ethyl or C 1-6 alkyl; or

(iii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, W is 3-cyclopropylaziridin-2-yl or optionally substituted aziridine and R L1 and R L2 taken together form fused cyclopropyl, then R 2 is not ethyl or C 1-6 alkyl; or

(iv) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene and two R L1 substituents taken together form spiro cyclopropyl, then W is not 3-cyclopropylaziridin-2-yl or optionally substituted aziridine; or

(v) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene and R L1 and R L2 taken together form fused cyclopropyl, then W is not 3-cyclopropylaziridin-2-yl or optionally substituted aziridine; or

(vi) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, W is 3-cyclopropylaziridin-2-yl or optionally substituted aziridine and two R L1 substituents taken together form spiro cyclopropyl, then L 1 is not —N(CH 3 )C(O)CH 2 N(CH 3 )C(O)—; or

(vii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, W is 3-cyclopropylaziridin-2-yl or optionally substituted aziridine and R L1 and R L2 taken together form fused cyclopropyl, then L 1 is not —N(CH 3 )C(O)CH 2 N(CH 3 )C(O)—.

4 . A compound, or a pharmaceutically acceptable salt thereof, having the structure of Formula IIc:

wherein the dotted lines represent zero, one, two, three, or four non-adjacent double bonds;

A is optionally substituted C 2 -C 4 alkylene, optionally substituted C 1 -C 4 heteroalkylene, optionally substituted C 2 -C 4 alkenylene, optionally substituted 3- to 6-membered cycloalkylene, optionally substituted 3- to 6-membered heterocycloalkylene, optionally substituted 6-membered arylene, or optionally substituted 5- to 10-membered heteroarylene;

B is absent, —CH(R 9 )—, >C═CR 9 R 9′ , or >CR 9 R 9′ where the carbon is bound to the carbonyl carbon of —N(R 11 )C(O)—, optionally substituted 3- to 6-membered cycloalkylene, optionally substituted 3- to 6-membered heterocycloalkylene, optionally substituted 6-membered arylene, or optionally substituted 5- to 6-membered heteroarylene;

G is optionally substituted C 1 -C 4 alkylene, optionally substituted C 1 -C 4 alkenylene, optionally substituted C 1 -C 4 heteroalkylene, —C(O)O—CH(R 6 )— where C is bound to —C(R 7 R 8 )—, —C(O)NH—CH(R 6 )—where C is bound to —C(R 7 R 8 )—, optionally substituted C 1 -C 4 heteroalkylene, or 3- to 8-membered heteroarylene;

L 1 is a linker;

W is a cross-linking group comprising a vinyl ketone, a vinyl sulfone, an ynone, a haloacetyl, or an alkynyl sulfone;

L has the structure of Formula VIIa or VIIb:

z is 0, 1, or 2;

X 9 is —NR L6 —, —C(O)—, or —S(O) 2 —;

each of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 is, independently, hydrogen, halogen, hydroxyl, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, or optionally substituted C 1 -C 6 heteroalkyl; or any two of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 together with the atoms to which they are attached and any intervening atoms to form an optionally substituted C 3 -C 8 cycloalkyl or a 3- to 8-membered heterocyclyl;

wherein L does not have the structure of

 or

optionally, wherein L is not

X 1 is optionally substituted C 1 -C 2 alkylene, NR, O, or S(O) q ;

X 2 is O or NH;

X 3 is N or CH;

q is 0, 1, or 2;

R is hydrogen, cyano, optionally substituted C 1 -C 4 alkyl, optionally substituted C 2 -C 4 alkenyl, optionally substituted C 2 -C 4 alkynyl, C(O)R′, C(O)OR′, C(O)N(R′) 2 , S(O)R′, S(O) 2 R′, or S(O) 2 N(R′) 2 ;

each R′ is, independently, hydrogen or optionally substituted C 1 -C 4 alkyl;

Y 1 is C, CH, or N;

Y 2 , Y 3 , Y 4 , and Y 7 are, independently, C or N;

Y 5 is CH, CH 2 , or N;

Y 6 is C(O), CH, CH 2 , or N;

R 13 is cyano, optionally substituted C 1 -C 6 alkyl, optionally substituted C 1 -C 6 heteroalkyl, optionally substituted 3- to 6-membered cycloalkyl, optionally substituted 3- to 6-membered cycloalkenyl, optionally substituted 3- to 6-membered heterocycloalkyl, optionally substituted 6- to 10-membered aryl, or optionally substituted 5- to 10-membered heteroaryl, or

R 13 and R 2 combine with the atoms to which they are attached to form an optionally substituted 3- to 14-membered heterocycloalkyl;

R 2 is absent, hydrogen, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3- to 6-membered cycloalkyl, optionally substituted 3- to 7-membered heterocycloalkyl, optionally substituted 6-membered aryl, optionally substituted 5- or 6-membered heteroaryl; R 14 is absent or R 2 and R 14 combine with the atom to which they are attached to form an optionally substituted 3- to 8-membered cycloalkyl or optionally substituted 3- to 14-membered heterocycloalkyl;

R 15 is absent, hydrogen, halogen, cyano, or methyl optionally substituted with 1 to 3 halogens;

R 5 is hydrogen, C 1 -C 4 alkyl optionally substituted with halogen, cyano, hydroxy, or C 1 -C 4 heteroalkyl, cyclopropyl, or cyclobutyl;

R 6 is hydrogen or methyl; R 7 is hydrogen, halogen, or optionally substituted C 1 -C 3 alkyl, or

R 6 and R 7 combine with the carbon atoms to which they are attached to form an optionally substituted 3- to 6-membered cycloalkyl or optionally substituted 3- to 7-membered heterocycloalkyl;

R 8 is hydrogen, halogen, hydroxy, cyano, optionally substituted C 1 -C 3 heteroalkyl, optionally substituted C 1 -C 3 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3- to 8-membered cycloalkyl, optionally substituted 3- to 14-membered heterocycloalkyl, optionally substituted 5- to 10-membered heteroaryl, or optionally substituted 6- to 10-membered aryl, or

R 7 and R 8 combine with the carbon atom to which they are attached to form C═CR 7′ R 8′ ; C═N(OH), C═N(O—C 1 -C 3 alkyl), C═O, C═S, C═NH, optionally substituted 3- to 6-membered cycloalkyl, or optionally substituted 3- to 7-membered heterocycloalkyl;

R 7a and R 8a are, independently, hydrogen, halogen, optionally substituted C 1 -C 3 alkyl, or combine with the carbon to which they are attached to form a carbonyl;

R 7′ is hydrogen, halogen, or optionally substituted C 1 -C 3 alkyl; R 8′ is hydrogen, halogen, hydroxy, cyano, optionally substituted C 1 -C 3 heteroalkyl, optionally substituted C 1 -C 3 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3- to 8-membered cycloalkyl, optionally substituted 3- to 14-membered heterocycloalkyl, optionally substituted 5- to 10-membered heteroaryl, or optionally substituted 6- to 10-membered aryl, or

R 7′ and R 8′ combine with the carbon atom to which they are attached to form optionally substituted 3- to 6-membered cycloalkyl or optionally substituted 3- to 7-membered heterocycloalkyl;

R 9 is hydrogen, fluoro, optionally substituted C 1 -C 6 alkyl, optionally substituted C 1 -C 6 heteroalkyl, optionally substituted 3- to 6-membered cycloalkyl, or optionally substituted 3- to 7-membered heterocycloalkyl; or

R 9 and L combine with the atoms to which they are attached to form an optionally substituted 3- to 14-membered heterocycloalkyl;

R 9′ is hydrogen or optionally substituted C 1 -C 6 alkyl; or

R 9 and R 9′ , combined with the atoms to which they are attached, form a 3- to 6-membered cycloalkyl or a 3- to 6-membered heterocycloalkyl;

R 10 is hydrogen, halogen, hydroxy, optionally substituted C 1 -C 3 heteroalkyl, or optionally substituted C 1 -C 3 alkyl;

R 10a is hydrogen or halogen;

R 11 is hydrogen or optionally substituted C 1 -C 3 alkyl; and

R 21 is hydrogen or optionally substituted C 1 -C 3 alkyl; wherein:

(i) the compound is not:

 or

(ii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, two R L1 substituents taken together form spiro cyclopropyl, and W is vinyl ketone, then R 2 is not ethyl or C 1-6 alkyl; or

(iii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, R L1 and R L2 taken together form fused cyclopropyl, and W is vinyl ketone, then R 2 is not ethyl or C 1-6 alkyl; or

(iv) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene and two R L1 substituents taken together form spiro cyclopropyl, then W is not vinyl ketone; or

(v) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, R L1 and R L2 taken together form fused cyclopropyl, then W is not vinyl ketone; or

(vi) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, W is vinyl ketone and two R L1 substituents taken together form spiro cyclopropyl or spiro cycloalkyl, then L 1 is other than

 or

(vii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, W is vinyl ketone and R L1 and R L2 taken together form fused cyclopropyl or fused cycloalkyl, then L 1 is other than

 or

(vii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, W is vinyl ketone and two R L1 substituents taken together form spiro cycloalkyl, then L 1 is not

 optionally substituted with 1, 2 or 3 independently selected C 1-6 alkyl; or

(ix) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, W is vinyl ketone and R L1 and R L2 taken together form fused cycloalkyl, then L 1 is not

 optionally substituted with 1, 2 or 3 independently selected C 1-6 alkyl.

5 . A compound, or a pharmaceutically acceptable salt thereof, having the structure of Formula III:

wherein A is optionally substituted 3- to 6-membered cycloalkylene, optionally substituted 3- to 6-membered heterocycloalkylene, optionally substituted 6-membered arylene, optionally substituted 5- to 6-membered heteroarylene, optionally substituted C 2 -C 4 alkylene, optionally substituted C 1 -C 4 heteroalkylene or optionally substituted C 2 -C 4 alkenylene;

Y 8 is

W is hydrogen, C 1 -C 4 alkyl, optionally substituted C 1 -C 3 heteroalkyl, optionally substituted 3- to 10-membered heterocycloalkyl, optionally substituted 3- to 10-membered cycloalkyl, optionally substituted 6- to 10-membered aryl, or optionally substituted 5- to 10-membered heteroaryl;

L has the structure of Formula VIIa or VIIb:

z is 0, 1, or 2;

X 9 is —NR L6 —, —C(O)—, or —S(O) 2 —;

each of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 is, independently, hydrogen, halogen, hydroxyl, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, or optionally substituted C 1 -C 6 heteroalkyl; or any two of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 together with the atoms to which they are attached and any intervening atoms to form an optionally substituted C 3 -C 8 cycloalkyl or a 3- to 8-membered heterocyclyl; or R L1 and R L2 taken together form a bond;

wherein L does not have the structure of

 or

optionally, wherein L is not

X 4 and X 5 are each, independently, CH 2 , CH(CH 3 ) or NH;

R 13 is optionally substituted C 1 -C 6 alkyl, optionally substituted C 1 -C 6 heteroalkyl, optionally substituted 3- to 6-membered cycloalkyl, optionally substituted 3- to 6-membered cycloalkenyl, optionally substituted 3- to 15-membered heterocycloalkyl, optionally substituted 6- to 10-membered aryl, or optionally substituted 5- to 10-membered heteroaryl;

R 2 is hydrogen, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3- to 6-membered cycloalkyl, optionally substituted 3- to 7-membered heterocycloalkyl, optionally substituted 6-membered aryl, optionally substituted 5- or 6-membered heteroaryl;

R 10 is hydrogen, hydroxy, optionally substituted C 1 -C 6 alkoxy, optionally substituted C 1 -C 3 alkyl, optionally substituted C 1 -C 6 heteroalkyl, or optionally substituted 3- to 10-membered heterocycloalkyl; and

R 7 and R 8 are each, independently, selected from fluoro or CH 3 , or R 7 and R 8 combine with the atoms to which they are attached to make a 3-membered cycloalkyl; wherein:

(i) the compound is not E10 to E52 in Table 2 and those set forth in Table 5; or

(ii) when A is thiazole-2,4-diyl and two R L1 substituents taken together form spiro cyclopropyl, then R 2 is not optionally substituted C 1-6 alkyl; or

(iii) when A is thiazole-2,4-diyl and two R L2 substituents taken together form spiro cyclopropyl, then R 2 is not optionally substituted C 1-6 alkyl; or

(iv) when A is thiazole-2,4-diyl and R L1 and R L2 taken together form optionally substituted fused cyclopropyl, then R 2 is not optionally substituted C 1-6 alkyl; or

(v) when A is thiazole-2,4-diyl and R L1 and R L2 taken together form a bond, then R 2 is not optionally substituted C 1-6 alkyl; or

(vi) when A is thiazole-2,4-diyl and R L1 and R L2 taken together form fused cyclobutyl, then R 2 is not optionally substituted C 1-6 alkyl; or

(vii) when A is thiazole-2,4-diyl and R L1 and R L3 together with the atoms to which they are attached form bridged cyclobutyl, then R 2 is not optionally substituted C 1-6 alkyl; or

(viii) when A is thiazole-2,4-diyl and R L1 and R L3 together with the atoms to which they are attached form bridged cyclobutyl, then W is not optionally substituted cyclopropyl.

6 . A compound, or pharmaceutically acceptable salt thereof, having the structure of Formula IV:

wherein A is optionally substituted 3- to 6-membered heterocycloalkylene, optionally substituted 3- to 6-membered cycloalkylene, optionally substituted 6-membered arylene, or optionally substituted 5- to 10-membered heteroarylene;

L 1 is a linker;

L has the structure of Formula VIIa or VIIb:

z is 0, 1, or 2;

X 9 is —NR L6 —, —C(O)—, or —S(O) 2 —;

each of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 is, independently, hydrogen, halogen, hydroxyl, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, or optionally substituted C 1 -C 6 heteroalkyl; or any two of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 together with the atoms to which they are attached and any intervening atoms to form an optionally substituted C 3 -C 8 cycloalkyl or a 3- to 8-membered heterocyclyl;

wherein L does not have the structure of

 or

optionally, wherein L is not

W is a cross-linking group comprising a vinyl ketone, vinyl sulfone, ynone, or an alkynyl sulfone;

R 1 is hydrogen, optionally substituted 3- to 10-membered heterocycloalkyl, or optionally substituted C 1 -C 6 heteroalkyl;

R 2 is optionally substituted C 1 -C 6 alkyl; and

R 3 is optionally substituted C 1 -C 6 alkyl or optionally substituted C 1 -C 3 heteroalkyl;

(i) the compound is not:

 or

(ii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, two R L1 substituents taken together form spiro cyclopropyl, and W is vinyl ketone, then R 2 is not ethyl; or

(iii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, R L1 and R L2 taken together form fused cyclopropyl, and W is vinyl ketone, then R 2 is not ethyl; or

(iv) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene and two R L1 substituents taken together form spiro cyclopropyl, then W is not vinyl ketone; or

(v) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, R L1 and R L2 taken together form fused cyclopropyl, then W is not vinyl ketone; or

(vi) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, W is vinyl ketone and two R L1 substituents taken together form spiro cyclopropyl or spiro cycloalkyl, then L 1 is other than

 or

(vii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, W is vinyl ketone and R L1 and R L2 taken together form fused cyclopropyl or fused cycloalkyl, then L 1 is not

 or

(viii) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, W is vinyl ketone and two R L1 substituents taken together form spiro cyclopropyl or spiro cycloalkyl, then L 1 is not

 optionally substituted with 1, 2 or 3 independently selected C 1-6 alkyl; or

(ix) when A is 1-hydroxy-benzene-3,5-diyl or optionally substituted phenylene, W is vinyl ketone and two R L1 and R L2 taken together form fused cyclopropyl or fused cycloalkyl, then L 1 is not

 optionally substituted with 1, 2 or 3 independently selected C 1-6 alkyl.

7 . A compound, or a pharmaceutically acceptable salt thereof, having the structure of Formula V:

wherein A is optionally substituted 3- to 6-membered heterocycloalkylene, optionally substituted 3- to 6-membered cycloalkylene, optionally substituted 6-membered arylene, or optionally substituted 5- to 10-membered heteroarylene;

L has the structure of Formula VIIa or VIIb:

z is 0, 1, or 2;

X 9 is —NR L6 —, —C(O)—, or —S(O) 2 —;

each of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 is, independently, hydrogen, halogen, hydroxyl, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, or optionally substituted C 1 -C 6 heteroalkyl; or any two of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 together with the atoms to which they are attached and any intervening atoms to form an optionally substituted C 3 -C 8 cycloalkyl or a 3- to 8-membered heterocyclyl;

wherein L does not have the structure of

 or

optionally, wherein L is not

W is a cross-linking group comprising an aziridine, an epoxide, a carbodiimide, an oxazoline, a thiazoline, a chloroethyl urea, a chloroethyl thiourea, a chloroethyl carbamate, a chloroethyl thiocarbamate, a trifluoromethyl ketone, a boronic acid, a boronic ester, an N-ethoxycarbonyl-2-ethoxy-1,2-dihydroquinoline (EEDQ), an iso-EEDQ or other EEDQ derivative, an oxazolium, or a glycal;

X 6 is CH 2 or O;

m is 1 or 2;

n is 0 or 1;

R 1 is hydrogen or optionally substituted 3- to 10-membered heterocycloalkyl; and

R 2 is optionally substituted C 1 -C 6 alkyl.

8 . A compound, or a pharmaceutically acceptable salt thereof, having the structure of Formula VI:

wherein A is optionally substituted 3- to 6-membered heterocycloalkylene, optionally substituted 3- to 6-membered cycloalkylene, optionally substituted 6-membered arylene, or optionally substituted 5- to 10-membered heteroarylene;

L has the structure of Formula VIIa or VIIb:

optionally, wherein L is not

z is 0, 1, or 2;

X 9 is —NR L6 —, —C(O)—, or —S(O) 2 —;

each of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 is, independently, hydrogen, halogen, hydroxyl, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, or optionally substituted C 1 -C 6 heteroalkyl; or any two of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 together with the atoms to which they are attached and any intervening atoms to form an optionally substituted C 3 -C 8 cycloalkyl or a 3- to 8-membered heterocyclyl;

X 6 , X 7 , and X 8 are each independently selected from CH 2 , CHF, CF 2 , C═O, or O;

m is 1 or 2;

n is 0 or 1;

R 1 is hydrogen, optionally substituted C 1 -C 6 heteroalkyl, or optionally substituted 3- to 10-membered heterocycloalkyl;

R 2 is optionally substituted C 1 -C 6 alkyl; and

R 3 is optionally substituted C 1 -C 6 alkyl, optionally substituted C 1 -C 6 heteroalkyl, optionally substituted 3- to 6-membered cycloalkyl, or optionally substituted heterocycloalkyl,

and wherein each hydrogen is independently, optionally, isotopically enriched for deuterium.

9 . A compound having Formula VIIIa:

or a pharmaceutically acceptable salt thereof, wherein:

the dotted lines represent zero, one, two, three, or four non-adjacent double bonds;

L has the structure of Formula VIIa or VIIb:

z is 0, 1, or 2;

X 9 is —NR L6 —, —C(O)—, or —S(O) 2 —;

each of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 is, independently, hydrogen, halogen, hydroxyl, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted C 1 -C 6 heteroalkyl, or optionally substituted C 3 -C 8 cycloalkyl; or any two of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 together with the atoms to which they are attached and any intervening atoms to form an optionally substituted C 3 -C 8 cycloalkyl or a 3- to 8-membered heterocyclyl;

wherein L does not have the structure of

 or

optionally, wherein L is not

A is optionally substituted C 2 -C 4 alkylene, optionally substituted C 1 -C 4 heteroalkylene, or optionally substituted C 2 -C 4 alkenylene;

G is optionally substituted C 1 -C 4 alkylene, optionally substituted C 1 -C 4 alkenylene, optionally substituted C 1 -C 4 heteroalkylene, —C(O)O—CH(R 6 )— where C is bound to —C(R 7 R 8 )—, —C(O)NH—CH(R 6 )—where C is bound to —C(R 7 R 8 )—, optionally substituted C 1 -C 4 heteroalkylene, or 3 to 8-membered heteroarylene;

swIp (Switch I/P-loop) is an organic moiety that non-covalently binds to both the Switch I binding pocket and residues 12 or 13 of the P-loop of a Ras protein;

X 3 is N or CH;

Y 1 is C, CH, or N;

Y 2 , Y 3 , Y 4 , and Y 7 are, independently, C or N;

Y 5 is CH, CH 2 , or N;

Y 6 is C(O), CH, CH 2 , or N;

R 1 is cyano, optionally substituted C 1 -C 6 alkyl, optionally substituted C 1 -C 6 heteroalkyl, optionally substituted 3 to 6-membered cycloalkyl, optionally substituted 3 to 6-membered cycloalkenyl, optionally substituted 3 to 6-membered heterocycloalkyl, optionally substituted 6 to 10-membered aryl, or optionally substituted 5 to 10-membered heteroaryl, or

R 1 and R 2 combine with the atoms to which they are attached to form an optionally substituted 3 to 14-membered heterocycloalkyl;

R 2 is absent, hydrogen, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3 to 6-membered cycloalkyl, optionally substituted 3 to 7-membered heterocycloalkyl, optionally substituted 6-membered aryl, optionally substituted 5 or 6-membered heteroaryl;

R 3 is absent, or

R 2 and R 3 combine with the atom to which they are attached to form an optionally substituted 3 to 8-membered cycloalkyl or optionally substituted 3 to 14-membered heterocycloalkyl;

R 4 is absent, hydrogen, halogen, cyano, or methyl optionally substituted with 1 to 3 halogens;

R 6 is hydrogen or methyl; R 7 is hydrogen, halogen, or optionally substituted C 1 -C 3 alkyl, or

R 6 and R 7 combine with the carbon atoms to which they are attached to form an optionally substituted 3 to 6-membered cycloalkyl or optionally substituted 3 to 7-membered heterocycloalkyl;

R 8 is hydrogen, halogen, hydroxy, cyano, optionally substituted C 1 -C 3 alkoxy, optionally substituted C 1 -C 3 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3 to 8-membered cycloalkyl, optionally substituted 3 to 14-membered heterocycloalkyl, optionally substituted 5 to 10-membered heteroaryl, or optionally substituted 6 to 10-membered aryl, or

R 7 and R 8 combine with the carbon atom to which they are attached to form C═CR 7′ R 8′ ; C═N(OH), C═N(O—C 1 -C 3 alkyl), C═O, C═S, C═NH, optionally substituted 3 to 6-membered cycloalkyl, or optionally substituted 3 to 7-membered heterocycloalkyl;

R 7a and R 8a are, independently, hydrogen, halo, optionally substituted C 1 -C 3 alkyl, or combine with the carbon to which they are attached to form a carbonyl;

R 7′ is hydrogen, halogen, or optionally substituted C 1 -C 3 alkyl; R 8′ is hydrogen, halogen, hydroxy, cyano, optionally substituted C 1 -C 3 alkoxy, optionally substituted C 1 -C 3 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3 to 8-membered cycloalkyl, optionally substituted 3 to 14-membered heterocycloalkyl, optionally substituted 5 to 10-membered heteroaryl, or optionally substituted 6 to 10-membered aryl, or

R 7′ and R 8′ combine with the carbon atom to which they are attached to form optionally substituted 3 to 6-membered cycloalkyl or optionally substituted 3 to 7-membered heterocycloalkyl;

R 10 is hydrogen, halo, hydroxy, C 1 -C 3 alkoxy, or C 1 -C 3 alkyl; and

R 10a is hydrogen or halo.

10 . The compound of claim 9 , or a pharmaceutically acceptable salt thereof, wherein the compound has formula VIIIb, VIIIc, VIIId, VIIIe, VIIIf or VIIIg:

wherein at each occurrence,

each L is independently having Formula VIIa or VIIb:

z is 0, 1, or 2;

X 9 is —NR L6 —, —C(O)—, or —S(O) 2 —;

each of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 is, independently, hydrogen, halogen, hydroxyl, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted C 1 -C 6 heteroalkyl, or optionally substituted C 3 -C 8 cycloalkyl; or any two of R L1 , R L2 , R L3 , R L4 , R L4 , R L5 , and R L6 together with the atoms to which they are attached and any intervening atoms to form an optionally substituted C 3 -C 8 cycloalkyl or a 3- to 8-membered heterocyclyl;

wherein L does not have the structure of

G is optionally substituted C 1 -C 4 alkylene, optionally substituted C 1 -C 4 alkenylene, optionally substituted C 1 -C 4 heteroalkylene, —C(O)O—CH(R 6 )— where C is bound to —C(R 7 R 8 )—, —C(O)NH—CH(R 6 )—where C is bound to —C(R 7 R 8 )—, optionally substituted C 1 -C 4 heteroalkylene, or 3 to 8-membered heteroarylene;

Z is —C(O)— or —S(O) 2 —;

each B is independently absent, —NH—, —N(CH 3 )—, —O—, —CH(R 9 )— or >C═CR 9 R 9′ where the carbon is bound to the carbonyl carbon of —N(R 11 )C(O)—, optionally substituted 3 to 6-membered cycloalkylene, optionally substituted 3 to 6-membered heterocycloalkylene, optionally substituted 6-membered arylene, or 5 to 6-membered heteroarylene;

each L 1 is independently absent or a linker;

each W is independently hydrogen, cyano, optionally substituted amino, optionally substituted amido, optionally substituted C 1 -C 4 alkoxy, optionally substituted C 1 -C 4 hydroxyalkyl, optionally substituted C 1 -C 4 aminoalkyl, optionally substituted C 1 -C 4 haloalkyl, optionally substituted C 1 -C 4 alkyl, optionally substituted C 1 -C 4 guanidinoalkyl, C 0 -C 4 alkyl optionally substituted 3 to 11-membered heterocycloalkyl, optionally substituted 3 to 10-membered cycloalkyl, optionally substituted 6 to 10-membered aryl, or optionally substituted 3 to 10-membered heteroaryl;

each X 3 is independently N or CH;

X e is N, CH, or CR 17 ;

X f is N or CH;

R 12 is optionally substituted C 1 -C 6 alkyl or optionally substituted C 1 -C 6 heteroalkyl;

R 17 is optionally substituted C 1 -C 6 alkyl, optionally substituted C 1 -C 6 heteroalkyl, optionally substituted 3 to 6-membered cycloalkyl, optionally substituted 3 to 6-membered cycloalkenyl, optionally substituted 3 to 6-membered heterocycloalkyl, optionally substituted 6 to 10-membered aryl, or optionally substituted 5 to 10-membered heteroaryl.

each Y 1 is independently C, CH, or N;

Y 2 , Y 3 , Y 4 , and Y 7 are, independently, C or N;

each Y 5 is independently CH, CH 2 , or N;

each Y 6 is independently C(O), CH, CH 2 , or N;

each R 1 is independently cyano, optionally substituted C 1 -C 6 alkyl, optionally substituted C 1 -C 6 heteroalkyl, optionally substituted 3 to 6-membered cycloalkyl, optionally substituted 3 to 6-membered cycloalkenyl, optionally substituted 3 to 6-membered heterocycloalkyl, optionally substituted 6 to 10-membered aryl, or optionally substituted 5 to 10-membered heteroaryl, or

R 1 and R 2 combine with the atoms to which they are attached to form an optionally substituted 3 to 14-membered heterocycloalkyl;

each R 2 is independently absent, hydrogen, optionally substituted C 1 -C 6 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3 to 6-membered cycloalkyl, optionally substituted 3 to 7-membered heterocycloalkyl, optionally substituted 6-membered aryl, optionally substituted 5 or 6-membered heteroaryl;

each R 3 is independently absent, or

R 2 and R 3 combine with the atom to which they are attached to form an optionally substituted 3 to 8-membered cycloalkyl or optionally substituted 3 to 14-membered heterocycloalkyl;

each R 4 is independently absent, hydrogen, halogen, cyano, or methyl optionally substituted with 1 to 3 halogens;

each R 6 is independently hydrogen or methyl; each R 7 is independently hydrogen, halogen, or optionally substituted C 1 -C 3 alkyl, or

R 6 and R 7 combine with the carbon atoms to which they are attached to form an optionally substituted 3 to 6-membered cycloalkyl or optionally substituted 3 to 7-membered heterocycloalkyl;

each R 8 is independently hydrogen, halogen, hydroxy, cyano, optionally substituted C 1 -C 3 alkoxy, optionally substituted C 1 -C 3 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3 to 8-membered cycloalkyl, optionally substituted 3 to 14-membered heterocycloalkyl, optionally substituted 5 to 10-membered heteroaryl, or optionally substituted 6 to 10-membered aryl, or

R 7 and R 8 combine with the carbon atom to which they are attached to form C═CR 7′ R 8′ ; C═N(OH), C═N(O—C 1 -C 3 alkyl), C═O, C═S, C═NH, optionally substituted 3 to 6-membered cycloalkyl, or optionally substituted 3 to 7-membered heterocycloalkyl;

R 7a and R 8a are, independently, hydrogen, halo, optionally substituted C 1 -C 3 alkyl, or combine with the carbon to which they are attached to form a carbonyl;

R 7′ is hydrogen, halogen, or optionally substituted C 1 -C 3 alkyl; R 8′ is hydrogen, halogen, hydroxy, cyano, optionally substituted C 1 -C 3 alkoxy, optionally substituted C 1 -C 3 alkyl, optionally substituted C 2 -C 6 alkenyl, optionally substituted C 2 -C 6 alkynyl, optionally substituted 3 to 8-membered cycloalkyl, optionally substituted 3 to 14-membered heterocycloalkyl, optionally substituted 5 to 10-membered heteroaryl, or optionally substituted 6 to 10-membered aryl, or

R 7′ and R 8′ combine with the carbon atom to which they are attached to form optionally substituted 3 to 6-membered cycloalkyl or optionally substituted 3 to 7-membered heterocycloalkyl;

each R 9 is independently hydrogen, F, optionally substituted C 1 -C 6 alkyl, optionally substituted C 1 -C 6 heteroalkyl, optionally substituted 3 to 6-membered cycloalkyl, or optionally substituted 3 to 7-membered heterocycloalkyl;

R 9 and L combine with the atoms to which they are attached to form an optionally substituted 3 to 14-membered heterocycloalkyl;

each R 9′ is independently hydrogen or optionally substituted C 1 -C 6 alkyl;

each R 10 is independently hydrogen, halo, hydroxy, C 1 -C 3 alkoxy, or C 1 -C 3 alkyl;

R 10a is hydrogen or halo;

R 11 is hydrogen or C 1 -C 3 alkyl.

11 . The compound of any one of claims 1 to 10 , or a pharmaceutically acceptable salt thereof, wherein L has the structure of Formula VIIa, Formula VIIa-1, Formula VIIa-2, Formula VIIa-3, Formula VIIa-4, Formula VIIa-5, Formula VIIa-6, Formula VIIb, Formula VIIb-1, Formula VIIb-2, or

wherein X 9 is —NR L6 —; or X 9 is —C(O)—; or X 9 is S(O) 2 —.

12 . The compound of any one of claims 1 to 11 , or a pharmaceutically acceptable salt thereof, wherein z is 0; or z is 1; or z is 2.

13 . The compound of any one of claims 1 to 12 , or a pharmaceutically acceptable salt thereof, wherein R L1 is hydrogen; or R L1 is optionally substituted C 1 -C 6 alkyl; or R L1 is methyl, ethyl, or trifluoromethyl; or R L1 is optionally substituted C 1 -C 6 heteroalkyl; or R L1 is methoxy or ethoxy; R L1 is optionally substituted C 2 -C 6 alkynyl; or R L1 is ethynyl.

14 . The compound of any one of claims 1 to 13 , or a pharmaceutically acceptable salt thereof, wherein R L2 is hydrogen; or R L2 is halogen; or R L2 is fluoro.

15 . The compound of any one of claims 1 to 14 , or a pharmaceutically acceptable salt thereof, wherein R L3 is hydrogen; or R L3 is optionally substituted C 1 -C 6 alkyl; or R L3 is methyl.

16 . The compound of any one of claims 1 to 15 , or a pharmaceutically acceptable salt thereof, wherein:

R L4 is hydrogen; or

R L1 and R L4 combine to form an optionally substituted C 4 cycloalkyl; or

R L1 and R L3 combine to form an optionally substituted C 4 cycloalkyl; or

R L1 and R L3 combine to form an optionally substituted C 5 cycloalkyl; or

two R L1 combine to form an optionally substituted C 3 -C 6 cycloalkyl; or

R L1 and R L2 combine to form an optionally substituted C 3 -C 6 cycloalkyl.

17 . The compound of any one of claims 1 to 16 , or a pharmaceutically acceptable salt thereof, wherein L is:

 or

L is:

18 . The compound of any one of claims 1 to 17 , or a pharmaceutically acceptable salt thereof, wherein R L6 is optionally substituted C 1 -C 6 alkyl; or R L6 is methyl.

19 . The compound of any one of claims 1 to 18 , or a pharmaceutically acceptable salt thereof, wherein R L5 is hydrogen; or R L5 is optionally substituted C 1 -C 6 alkyl; or R L5 is optionally substituted C 3 -C 8 cycloalkyl; or two R L5 combine to form an optionally substituted C 3 -C 8 cycloalkyl.

20 . The compound of any one of claims 1 to 19 , or a pharmaceutically acceptable salt thereof, wherein:

A is optionally substituted 3- to 6-membered heterocycloalkylene, optionally substituted 6-membered arylene, or optionally substituted 5- to 10-membered heteroarylene; or

A is optionally substituted 6-membered arylene; or

A is

 or

A is optionally substituted 3- to 6-membered heterocycloalkylene;

A is

 or

A is optionally substituted 5- to 10-membered heteroarylene; or

A is

21 . The compound of any one of claims 2 to 20 , or a pharmaceutically acceptable salt thereof, wherein R 2 is ethyl or haloethyl.

22 . The compound of any one of claims 1 and 6 to 21 , or a pharmaceutically acceptable salt thereof, wherein R 3 is optionally substituted C 1 -C 6 alkyl.

23 . A compound, or a pharmaceutically acceptable salt thereof, of Table 1.

24 . A pharmaceutical composition comprising a compound, or a pharmaceutically acceptable salt thereof, of any one of claims 1 to 23 and a pharmaceutically acceptable excipient.

25 . A method of treating cancer in a subject in need thereof, the method comprising administering to the subject a therapeutically effective amount of a compound, or a pharmaceutically acceptable salt thereof, of any one of claims 1 to 23 or a pharmaceutical composition of claim 24 ,

optionally wherein the cancer is pancreatic cancer, colorectal cancer, non-small cell lung cancer, or endometrial cancer;

or optionally wherein the cancer comprises a Ras mutation; or

optionally wherein the method further comprises administering an additional anti-cancer therapy, optionally wherein the additional anti-cancer therapy is an EGFR inhibitor, a second Ras inhibitor, a SHP2 inhibitor, a SOS1 inhibitor, a Raf inhibitor, a MEK inhibitor, an ERK inhibitor, a PI3K inhibitor, a PTEN inhibitor, an AKT inhibitor, an mTORC1 inhibitor, a BRAF inhibitor, a PD-L1 inhibitor, a PD-1 inhibitor, a CDK4/6 inhibitor, a HER2 inhibitor, or a combination thereof.

26 . A method of treating a Ras protein-related disorder in a subject in need thereof, the method comprising administering to the subject a therapeutically effective amount of a compound of any one of claims 1 to 23 , or a pharmaceutically acceptable salt thereof, or a pharmaceutical composition of claim 24 , optionally wherein the Ras protein is K-Ras; or

optionally wherein the method further comprises administering an additional anti-cancer therapy, optionally wherein the additional anti-cancer therapy is an EGFR inhibitor, a second Ras inhibitor, a SHP2 inhibitor, a SOS1 inhibitor, a Raf inhibitor, a MEK inhibitor, an ERK inhibitor, a PI3K inhibitor, a PTEN inhibitor, an AKT inhibitor, an mTORC1 inhibitor, a BRAF inhibitor, a PD-L1 inhibitor, a PD-1 inhibitor, a CDK4/6 inhibitor, a HER2 inhibitor, or a combination thereof.

27 . A method of inhibiting a Ras protein in a cell, the method comprising contacting the cell with an effective amount of a compound of any one of claims 1 to 23 , or a pharmaceutically acceptable salt thereof, or a pharmaceutical composition of claim 24 ,

optionally wherein the Ras protein is K-Ras;

or optionally wherein the cell is a cancer cell, optionally wherein the cancer cell is a pancreatic cancer cell, a colorectal cancer cell, a non-small cell lung cancer cell, or an endometrial cancer cell; or

optionally wherein the method further comprises administering an additional anti-cancer therapy, optionally wherein the additional anti-cancer therapy is an EGFR inhibitor, a second Ras inhibitor, a SHP2 inhibitor, a SOS1 inhibitor, a Raf inhibitor, a MEK inhibitor, an ERK inhibitor, a PI3K inhibitor, a PTEN inhibitor, an AKT inhibitor, an mTORC1 inhibitor, a BRAF inhibitor, a PD-L1 inhibitor, a PD-1 inhibitor, a CDK4/6 inhibitor, a HER2 inhibitor, or a combination thereof.

Assignments (2)
ASSIGNMENT OF ASSIGNOR'S INTEREST Recorded Apr 2, 2026
From: KNOX, JOHN E.; KOLTUN, ELENA S.; PARSONS, DYLAN E.; TOMLINSON, AIDAN; BURNETT, G. LESLIE; CREGG, JAMES; GILL, ADRIAN L.; BUCKL, ANDREAS
To: REVOLUTION MEDICINES, INC.
Reel/Frame 074257/0845 →
SECURITY INTEREST Recorded Mar 24, 2026
From: REVOLUTION MEDICINES, INC.
To: WILMINGTON TRUST, NATIONAL ASSOCIATION, AS ADMINISTRATIVE AGENT
Reel/Frame 074172/0328 →