NR4A super-repressors and methods of use thereof
The present disclosure provides super-repressors capable of inhibiting the transcription of NR4A1, NR4A2, and NR4A3 target genes. The super-repressors can be used to enhance the effector functions of immune cells, e.g., for adoptive cell therapy. Methods of treating disorders using the modified immune cells are also provided.
1 . A method of treating cancer in a subject in need thereof comprising administering to the subject an effective amount of cells comprising a polynucleotide comprising a nucleic acid encoding a polypeptide comprising a DNA-binding domain that specifically binds to both the NBRE element and the NurRE element, wherein
(a) the polypeptide inhibits the activity of NR4A1, NR4A2, and NR4A3; or
(b) the polypeptide is capable of inhibiting transcription of a gene operably associated with both the NBRE element and the NurRE element.
2 . The method of claim 1 , wherein the polypeptide does not comprise a transcription activation domain.
3 . The method of claim 1 , wherein the cells are T cells.
4 . The method of claim 3 , wherein the cells further comprise an engineered immune receptor.
5 . A method of treating cancer in a subject in need thereof comprising administering to the subject an effective amount of cells comprising a polynucleotide comprising a nucleic acid encoding a polypeptide comprising a DNA-binding domain that specifically binds to both the NBRE element and the NurRE element, wherein the polypeptide does not comprise a transcription activation domain.
6 . The method of claim 5 , wherein the DNA-binding domain is a polypeptide comprising the amino acid sequence of the DNA-binding domain of NR4A1, NR4A2, or NR4A3.
7 . The method of claim 6 , wherein the DNA-binding domain comprises the amino acid sequence of the DNA-binding domain of NR4A3.
8 . The method of claim 7 , wherein the DNA-binding domain of NR4A3 is a polypeptide comprising the amino acid sequence of SEQ ID NO:4.
9 . The method of claim 5 , wherein the DNA-binding domain is a polypeptide comprising the amino acid sequence of any one of SEQ ID NOs: 4, 5, and 50-53.
10 . The method of claim 5 , wherein the DNA-binding domain comprises a TAL effector DNA binding domain, a zinc finger, or a DNA-binding domain from a meganuclease.
11 . The method of claim 5 , wherein the polypeptide further comprises the amino acid sequence of the ligand-binding domain (LBD) of NR4A1, NR4A2, or NR4A3.
12 . The method of claim 5 , wherein the polypeptide further comprises a dimerization domain, a transcriptional repressor domain, or a chromatin compaction domain.
13 . The method of claim 5 , wherein the polypeptide comprises the amino acid sequence of any one of SEQ ID NOs: 29-43.
14 . The method of claim 5 , wherein the polypeptide further comprises a Krüppel-associated box (KRAB) domain.
15 . The method of claim 5 , wherein the polypeptide further comprises NCOR1 or a fragment thereof.
16 . The method of claim 5 , wherein the DNA-binding domain is a polypeptide comprising the amino acid sequence of the DNA-binding domain of NR4A3.
17 . The method of claim 16 , wherein the DNA-binding domain of NR4A3 is a polypeptide comprising the amino acid sequence of SEQ ID NO:4.
18 . The method of claim 5 , wherein the cells are T cells.
19 . The method of claim 18 , wherein the cells further comprise an engineered immune receptor.